Based on the info given, the facts are currently these:
1) The sister is not liable and never has been and, apart from being an occupier, doesn't have an interest in the dwelling (currently).
2) The 16 year old, presumably, isn't directly liable as he is under 18.
3) Someone else, presumably, is currently being held liable (who?).
4) The 16 year old has claimed HB.
Based on all the above, the 16 year old is entirely within his rights to claim HB. He can properly be treated as liable to pay "rent" (or monies for use and occupation) under Regulation 8 of the HB Regulations 2006.
To cover all bases, I would get the 16 year old to submit an appeal. If the LA has not yet issued a formal notification of decision, they must - a claim has been made, so a decision must be made on the claim. At the same time, I would ALSO get the sister to submit a claim "just in case" - make it clear that is the basis of the claim.
I'm not sure about CTAX liabilities, so I'll leave that for someone else.
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